The story takes place during a conflict between the Krakozhia Dukedom and the Grania Republic. In the midst of the fighting, a savior appears to the Krakozhia Dukedom, and it is a high school student named Yano Souhei. Two female soldiers, First Lieutenant Ruto and Second Lieutenant Haruka, appear in tanks to intrude on Souhei's everyday life, followed by the enemy soldier Shachirofu, all of whom use firearms without hesitation at his house. (Source: ANN)

Following a catastrophic war against a race of giants known as the Zentradi, humanity has escaped towards the center of the galaxy aboard a fleet of colonial vessels called the Macross Frontier. As the extraterrestrial threat is left further and further behind, life on Macross Frontier proceeds as usual. In the year 2059, a young mecha pilot trainee named Alto Saotome and his colleagues are preparing to perform an accompanying routine for the famous singer Sheryl Nome, who has come to Macross Frontier for a concert. During the performance, a biomechanical alien species known as the Vajra make a sudden appearance, breaking through the defensive perimeter surrounding the vessel and crash-landing near the concert venue, plunging the entire city into chaos. As the concertgoers evacuate, a young girl named Ranka Lee is left behind and gets targeted by the Vajra, but she is saved at the last minute by Alto. Following these events, the Strategic Military Services program notes Alto's skill in battle, resulting in his recruitment to combat the new alien threat. [Written by MAL Rewrite]

Even at their colleagues' wedding, the sparkling members of Marukawa Publishing's Emerald Department stand out. Having been invited to assist in providing entertainment for the guests, Masamune Takano, Ritsu Onodera, Yoshiyuki Hatori, Shouta Kisa, Kanade Mino, as well as Takafumi Yokozawa enjoy the festivities as much as their work allows. As love blossoms between two individuals, marriage marks a momentous occasion fit for self-reflection. Whether they like it or not, the men of Marukawa need to consider where their own relationships are headed.

Koyama Yuzu is running an antique shop. Mayu, a cat god (nekogami), is living off Yuzu and leads an idle life playing games. Lots of other gods visit Mayu and enjoy merrymaking.